1116-52882 - Close-up of a young lion (Panthera leo) sitting in the grass, turning head, Laikipia, Kenya

1116-52883 - Portrait of a young, male lion (Panthera leo) standing on savanna, staring at the camera, licking his lips, Laikipia, Kenya

1116-52881 - Portrait of a male lion (Panthera leo) standing among bushes, looking mournful at the camera, Laikipia, Kenya

1116-52880 - Close-up of a Lioness (Panthera leo) walking through the low bushes in sunshine, Laikipia, Kenya

1116-52879 - Leopard (Panthera pardus) walking across sloping rockface in sunshine, Laikipia, Kenya

1116-52878 - Leopard (Panthera pardus) sits watching camera on sunlit rock, Kenya

1116-52877 - Leopard (Panthera pardus) lies on rock surrounded by branches, Kenya

1116-52838 - Panorama of African bush elephants (Loxodonta africana) standing in the water playing with each other with their trunks and tusks, face to face, in Chobe National Park, Chobe, North-West, Botswana

1116-52837 - African bush elephant (Loxodonta africana) in the water washing and lifting river grass with its trunk and tusks in Chobe National Park, Chobe, North-West, Botswana

1116-52836 - African bush elephant (Loxodonta africana) standing on the savannah throwing dust over itself at Segera, Segera, Laikipia, Kenya

1116-52835 - African bush elephant (Loxodonta africana) standing on the savannah while nursing young calf at Segera, Segera, Laikipia, Kenya

1116-51270 - A leopard (Panthera pardus) sits in the forked trunk of a tree. It has a brown, spotted coat and is looking straight at the camera. Shot with a Nikon D850 in Serengeti National Park; Tanzania

1116-51268 - A leopard (Panthera pardus) lies on the branch of a tree with its head up. It has black spots on its brown fur coat and is looking for prey. Shot with a Nikon D850 in the Masai Mara; Kenya

1116-51266 - A male lion (Panthera leo) shows the Flehmen response by opening his mouth wide as if he were roaring. He has a golden coat that glows in the warm early morning light. Shot with a Nikon D850 in Serengeti National Park; Tanzania

1116-51732 - A male leopard (Panthera pardus) stares at the camera. It has a brown, spotted coat, whiskers and a green eye, Masai Mara National Reserve; Kenya

1116-51267 - A leopard (Panthera pardus) walks towards the camera on a sandy track. It has black spots on its brown fur coat and is raising its paw to take the next step. Shot with a Nikon D850 in the Masai Mara; Kenya

1116-51907 - Lion (Panthera leo) stands looking ahead on stepping stones, Etosha National Park; Otavi, Oshikoto, Namibia

1116-51269 - A leopard (Panthera pardus) stands in a tree that is covered in lichen. It has black spots on its brown fur coat and is turning its head to look up. Shot with a Nikon D850 in the Masai Mara; Kenya

1116-51078 - A lion cub (Panthera leo) walks through long grass towards the camera. It is lifting its right paw and staring intently, Serengeti National Park; Mara Region, Tanzania

1116-51262 - A young male lion (Panthera leo) pokes his head above a rocky ledge under a blue sky. He has a short mane and is staring straight at the camera. Shot with a Nikon D850 in Serengeti National Park; Tanzania

1116-51951 - Close-up portrait of a lioness (Panthera leo) lying down in the golden long grass on the savanna in the Etosha National Park; Otavi, Oshikoto, Namibia

1116-51908 - Lion (Panthera leo) lies on stepping stones looking up, Etosha National Park; Otavi, Oshikoto, Namibia

1116-51264 - A lioness (Panthera leo) walks down a gravel airstrip next to her young cub. They both have golden coats, made to glow in the warm early morning light. Shot with a Nikon D850 in Serengeti National Park; Tanzania

1116-51731 - A young male lion (Panthera leo) pokes his head above a rocky ledge. He has a short mane and is staring straight at the camera, Serengeti National Park; Tanzania

1116-51278 - An African bush elephant (Loxodonta africana) is throwing dust over itself with its trunk on a bare earth slope with trees in the background under a clear sky. It has mud stains on its trunk, and the dust is exploding in a cloud against its wrinkled grey skin. Shot with a Nikon D810 in Chobe National Park; Botswana

1116-51265 - A lioness (Panthera leo) is about to slap a male lion with its paw after mating. They both have golden coats and are standing on a patch of burnt grass in the warm evening light. Shot with a Nikon D810 in the Serengeti; Tanzania

1116-49282 - Lion cub (Panthera Leo) sits on rock among bushes, Serengeti, Tanzania

1116-49380 - Three male lions (Panthera leo) feed on dead buffalo (Syncerus caffer), Serengeti, Tanzania

1116-49359 - Male lion (Panthera leo) eats buffalo as giraffe (Giraffa camelopardalis tippelskirchii) watches, Serengeti, Tanzania

1116-49356 - Male leopard (Panthera pardus) sits in branches eyeing camera, Serengeti, Tanzania

1116-49350 - Lion cub (Panthera leo) lies in grass looking left, Serengeti, Tanzania

1116-49351 - Lioness (Panthera leo) lies in dappled sunshine yawning widely, Serengeti, Tanzania

1116-49338 - Close-up of lion cub (Panthera leo) lying on twigs, Serengeti, Tanzania

1116-49256 - Lioness (Panthera leo) lies eyeing camera in leafy bushes, Serengeti, Tanzania

1116-49358 - Male lion (Panthera leo) bites Cape buffalo (Syncerus caffer) from behind with another lion nearby, Serengeti, Tanzania

1116-49254 - Male lion (Panthera leo) stalks Masai giraffe (Giraffa camelopardalis tippelskirchii) in savannah, Serengeti, Tanzania

1116-49308 - Slow pan of male lion (Panthera leo) walking left, Serengeti, Tanzania

1116-49360 - Male lion (Panthera leo) pushed down Cape buffalo (Syncerus caffer) from behind, Serengeti, Tanzania

1116-49357 - Male lion (Panthera leo) attacks Cape buffalo (Syncerus caffer) from behind, Serengeti, Tanzania

1116-49284 - Male lion (Panthera leo) lying in grass looking up, Serengeti, Tanzanai

1116-49298 - Close-up of male lion (Panthera leo) head and shoulders, Serengeti, Tanzania

1116-49361 - Male lion (Panthera leo) stretches in grass facing right, Serengeti, Tanzania

1116-49377 - Three male lions (Panthera leo) take down Cape buffalo (Syncerus caffer), Serengeti, Tanzania

1116-49337 - Close-up of lion cub (Panthera leo) lying on sticks, Serengeti, Tanzania

1116-49376 - Three male lions (Panthera leo) feed on dead buffalo (Syncerus caffer), Serengeti, Tanzania

1116-49257 - Male lion (Panthera leo) lies on a rock looking at the camera, Serengeti, Tanzania

1116-49275 - Tail of leopard (Panthera pardus) hangs down while climbing tree, Serengeti, Tanzania

1116-49285 - Male lion (Panthera leo) stands on rock beside tree, Serengeti, Tanzania

1116-49318 - Lion cub (Panthera leo) sits on rock by tree, Serengeti, Tanzania

1116-49281 - Lion cub (Panthera leo) nuzzles another as sibling watches, Serengeti, Tanzania

1116-49253 - Three lion cubs (Panthera leo) sitting on a rock looking out with two lionesses in the background at dusk, Serengeti, Tanzania

1116-50369 - Male leopard (Panthera pardus) stands in tree looking out, Cottar's 1920s Safari Camp, Maasai Mara National Reserve; Kenya

1116-50377 - Lion cub (Panthera leo) looks back climbing tree trunk, Grumeti Serengeti Tented Camp, Serengeti National Park; Tanzania

1116-50413 - Silhouette of lioness (Panthera leo) at sunrise crossing ridge, Grumeti Serengeti Tented Camp, Serengeti National Park; Tanzania

1116-50354 - Three lion cubs (Panthera leo) lie together by bush, Cottar's 1920s Safari Camp, Maasai Mara National Park; Kenya

1116-50400 - Close-up of lioness (Panthera leo) sitting with blurred background, Cottar's 1920s Safari Camp, Maasai Mara National Reserve; Kenya

1116-50390 - Close-up of lioness lying growling at cub, Grumeti Serengeti Tented Camp, Serengeti National Park; Tanzania

1116-50423 - Lioness (Panthera leo) lies drinking from pool by cub, Grumeti Serengeti Tented Camp, Serengeti National Park; Tanzania

1116-50360 - A young male lion (Panthera leo) pokes his head above a rocky ledge under a blue sky. He has a short mane and is staring straight at the camera. Klein's Camp, Serengeti National Park; Tanzania

1116-50375 - Lioness and cub (Panthera leo) drink from muddy pool, Cottar's 1920s Safari Camp, Maasai Mara National Reserve; Kenya

1116-50417 - A leopard (Panthera pardus) sits in the forked trunk of a tree. It has a brown, spotted coat and is looking straight at the camera, Serengeti National Park; Tanzania

1116-50391 - Close-up of lioness (Panthera leo) in tree looking down, Grumeti Serengeti Tented Camp, Serengeti National Park; Tanzania

1116-50416 - Leopard (Panthera pardus) lies on diagonal branch watching camera, Klein's Camp, Serengeti National Park; Tanzania

1116-50415 - Lioness (Panthera leo) lies in tree looking through branches, Grumeti Serengeti Tented Camp, Serengeti National Park; Tanzania

1116-50373 - Lioness (Panthera leo) lies in silhouette against dawn sky, Grumeti Serengeti Tented Camp, Serengeti National Park; Tanzania

1116-50366 - Male lion (Panthera leo) stands staring down from tree, Cottar's 1920s Safari Camp, Maasai Mara National Reserve; Kenya

1116-50392 - Close-up of lion cub (Panthera leo) yawning in grass, Grumeti Serengeti Tented Camp, Serengeti National Park; Tanzania

1116-50395 - Close-up of leopard (Panthera pardus) on branch facing right, Cottar's 1920s Safari Camp, Maasai Mara National Reserve; Kenya

1116-50424 - Lioness and cub (Panthera Leo) face camera near Impala (Aepyceros melampus) harem, Grumeti Serengeti Tent Camp, Serengeti National Park; Tanzania

1116-50330 - A male lion (Panthera leo) lies with its head bathed in the golden light of dawn. It has a big mane and is staring into the distance,Klein's Camp, Serengeti National Park; Tanzania

1116-50403 - Monochrome close-up of lion (Panthera leo) face in shadows, Klein's Camp, Serengeti National Park; Tanzania

1116-50393 - Close-up of lion cub (Panthera leo) staring on branch, Grumeti Serengeti Tented Camp, Serengeti National Park; Tanzania

1116-50421 - Two lionesses and three cubs (Panthera leo) lie in a tree, Grumeti Serengeti Tented Camp, Serengeti National Park; Tanzania

1116-50394 - Close-up of lion cub (Panthera leo) sitting in bushes, Grumeti Serengeti Tented Camp, Serengeti National Park; Tanzania

1116-50397 - Close-up of leopard (Panthera pardus) asleep on lichen-covered branch, Cottar's 1920s Safari Camp, Maasai Mara National Reserve; Kenya

1116-50404 - Monochrome close-up of half male leopard (Panthera pardus) face, Cottar's 1920s Safari Camp, Maasai Mara National Reserve; Kenya

1116-50418 - Close-up of leopard (Panthera pardus) sitting in long grass, Grumeti Serengeti Tented Camp, Serengeti National Park; Tanzania

1116-50388 - Close-up of lioness (Panthera leo) standing yawning in grass, Grumeti Serengeti Tented Camp, Serengeti National Park; Tanzania

1116-50396 - Close-up of leopard (Panthera pardus) facing camera on branch, Cottar's 1920s Safari Camp, Maasai Mara National Reserve; Kenya

1116-50420 - Two lionesses (Panthera leo) lie drinking from water hole, Grumeti Serengeti Tented Camp, Serengeti National Park; Tanzania

1116-50387 - Close-up of male leopard (Panthera pardus) head turned left, Cottar's 1920s Safari Camp, Maasai Mara National Reserve; Kenya

1116-50389 - Close-up of lioness (Panthera leo) on branch lying asleep, Grumeti Serengeti Tented Camp, Serengeti National Park; Tanzania

1116-50368 - Male lion (Panthera leo) stands behind lioness on rock, Klein's Camp, Serengeti National Park; Tanzania

1116-50414 - Lioness (Panthera leo) walks on rock under blue sky, Cottar's 1920s Safari Camp, Maasai Mara National Reserve; Kenya

1116-50367 - Male lion (Panthera leo) stands on branch looking around, Cottar's 1920s Safari Camp, Maasai Mara National Reserve; Kenya

1116-50374 - Lioness (Panthera leo) lies on kopje looking over rocks, Klein's Camp, Serengeti National Park; Tanzania

1116-50426 - Impala (Aepyceros melampus) harem watches lioness and cub (Panthera leo) approach, Grumeti Serengeti Tent Camp, Serengeti National Park; Tanzania

1116-50378 - Lion cub (Panthera leo) clutches tree trunk looking up, Grumeti Serengeti Tented Camp, Serengeti National Park; Tanzania

1116-50376 - Lion cubs (Panthera leo) look out from tree trunk, Grumeti Serengeti Tented Camp, Serengeti National Park; Tanzania

1116-47995 - A mother cheetah (Acinonyx jubatus) and her cub chase a scrub hare (Lepus saxatilis) on a grassy plain. They have golden fur covered with black spots, and the cub is stretching out it's paw to catch the hare. Masai Mara, Kenya

1116-48632 - Close-up of male lion (panthera Leo) alongside scratched tree, Serengeti National Park, Tanzania

1116-47517 - Male lion (Panthera leo) lies in grass staring left, Serengeti National Park, Tanzania

1116-47909 - Close-up of African buffalo (Syncerus caffer)head through grass, Maasai Mara National Reserve, Kenya

1116-47931 - Six blue wildebeest (Connochaetes taurinus) silhouetted against setting sun, Maasai Mara National Reserve, Kenya

1116-47518 - Head of male lion (Panthera leo) lying on kopje, Serengeti National Park, Tanzania

1116-47930 - Three blue wildebeest (Connochaetes taurinus) silhouetted against setting sun, Maasai Mara National Reserve, Kenya

1116-47522 - Close-up of male lion (Panthera leo) head staring out, Serengeti National Park, Tanzania

1116-47998 - A leopard (Panthera pardus) lies on the branch of a tree with it's head up. It has black spots on its brown fur coat and is looking for prey, Maasai Mara National Reserve, Kenya

1116-47264 - Leopard (Panthera pardus) crouching in tree in the Ndutu area of the Ngorongoro Crater Conservation Area on the Serengeti Plains, Tanzania

1116-47500 - Close-up of African elephant (Loxodonta africana) enjoying dust bath, Serengeti National Park, Tanzania