1116-52883 - Portrait of a young, male lion (Panthera leo) standing on savanna, staring at the camera, licking his lips, Laikipia, Kenya

1116-52882 - Close-up of a young lion (Panthera leo) sitting in the grass, turning head, Laikipia, Kenya

1116-52881 - Portrait of a male lion (Panthera leo) standing among bushes, looking mournful at the camera, Laikipia, Kenya

1116-52880 - Close-up of a Lioness (Panthera leo) walking through the low bushes in sunshine, Laikipia, Kenya

1116-52879 - Leopard (Panthera pardus) walking across sloping rockface in sunshine, Laikipia, Kenya

1116-52878 - Leopard (Panthera pardus) sits watching camera on sunlit rock, Kenya

1116-52877 - Leopard (Panthera pardus) lies on rock surrounded by branches, Kenya

1116-52872 - Close-up of Nile crocodile head (Crocodylus niloticus) on riverbank in Chobe National Park, Botswana

1116-52871 - Close-up of Nile crocodile foot (Crocodylus niloticus) in sunshine in Chobe National Park, Botswana

1116-52869 - Portrait of a Cape Buffalo (Syncerus caffer) drinking from river carrying two cattle egrets (Bubulcus ibis) on its back in Chobe National Park, Chobe, Botswana

1116-52870 - Mono, close-up portrait of a Cape Buffalo (Syncerus caffer) standing, eyeing camera in Segera, Segera, Laikipia, Kenya

1116-52868 - Portrait of a Cape Buffalo (Syncerus caffer) drinking from river carrying two cattle egrets (Bubulcus ibis) on its back in Chobe National Park, Chobe, Botswana

1116-52867 - Leopard tortoise (Stigmochelys pardalis) walks past over dry grass, Laikipia, Kenya

1116-52865 - Aldabra giant tortoise (Aldabrachelys gigantea) crosses lawn near trees, Segera, Laikipia, Kenya

1116-52866 - Close-up of Cape buffalo (Syncerus caffer) carrying Red-billed Oxpecker (Buphagus erythrorhynchus) in Chobe National Park, Botswana

1116-52864 - Cape buffalo (Syncerus caffer) walks through shallows watching camera in Chobe National Park, Botswana

1116-52863 - Young African elephant (Loxodonta africana) throwing sand over head in Chobe National Park, Botswana

1116-52861 - Three African bush elephants (Loxodonta africana) walk towards camera in Chobe National Park, Botswana

1116-52862 - Three Chacma baboons (Papio ursinus) sit on dead branch in Chobe National Park, Botswana

1116-52860 - Close-up of Vervet monkey (Chlorocebus pygerythrus) sitting grooming baby in Chobe National Park, Botswana

1116-52859 - Close-up of Chacma baboon (Papio ursinus) sitting holding baby in Chobe National Park, Botswana

1116-52858 - Close-up of baby Chacma baboon (Papio ursinus) drinking milk in Chobe National Park, Botswana

1116-52857 - Close-up of baby African elephant (Loxodonta africana) rolling trunk, Segera, Laikipia, Kenya

1116-52856 - Close-up of baboon (Papio ursinus) grooming another with baby in Chobe National Park, Botswana

1116-52855 - Close-up of African elephant (Loxodonta africana) feeding in river in Chobe National Park, Botswana

1116-52853 - Close-up of African bush elephant (Loxodonta africana) hind legs and tail, Segera, Laikipia, Kenya

1116-52854 - Close-up of African elephant (Loxodonta africana) eating in river in Chobe National Park, Botswana

1116-52852 - Baby African bush elephant (Loxodonta africana) walks between parents in Chobe National Park, Botswana

1116-52851 - Baby African bush elephant (Loxodonta africana) lying in mud in Chobe National Park, Botswana

1116-52849 - African bush elephant (Loxodonta africana) follows another in river in Chobe National Park, Botswana

1116-52850 - Baby African bush elephant (Loxodonta africana) crosses dirt track in Chobe National Park, Botswana

1116-52848 - African bush elephant (Loxodonta africana) flaps ears in river in Chobe National Park, Botswana

1116-52847 - African bush elephant (Loxodonta africana) confronts male Common waterbuck (Kobus ellipsiprymnus) in Chobe National Park, Botswana

1116-52846 - Close-up of dust exploding on African elephant (Loxodonta africana), Segera, Laikipia, Kenya

1116-52845 - African bush elephant (Loxodonta africana) stands splashing muddy water in Chobe National Park, Botswana

1116-52844 - African bush elephant (Loxodonta africana) nurses calf under tree, Kenya

1116-52843 - African bush elephant (Loxodonta africana) blows sand over head in Chobe National Park, Botswana

1116-52838 - Panorama of African bush elephants (Loxodonta africana) standing in the water playing with each other with their trunks and tusks, face to face, in Chobe National Park, Chobe, North-West, Botswana

1116-52837 - African bush elephant (Loxodonta africana) in the water washing and lifting river grass with its trunk and tusks in Chobe National Park, Chobe, North-West, Botswana

1116-52836 - African bush elephant (Loxodonta africana) standing on the savannah throwing dust over itself at Segera, Segera, Laikipia, Kenya

1116-52835 - African bush elephant (Loxodonta africana) standing on the savannah while nursing young calf at Segera, Segera, Laikipia, Kenya

1116-52803 - Three Southern giraffes (Giraffa giraffa angolensis) walk along sandy track in Chobe National Park, Botswana

1116-52801 - Reticulated giraffe (Giraffa camelopardalis reticulata) stands on dam turning head towards the camera, Segera, Laikipia, Kenya

1116-52802 - Reticulated giraffe (Giraffa camelopardalis reticulata) stands on dam watching camera, Segera, Laikipia, Kenya

1116-52800 - Reticulated giraffe (Giraffa camelopardalis reticulata) stands behind bush opening mouth, Segera, Laikipia, Kenya

1116-52799 - Close-up of Reticulated giraffe (Giraffa camelopardalis reticulata) gazing at camera, Segera, Laikipia, Kenya

1116-52780 - Reticulated giraffe (Giraffa reticulata) stands on the savannah browsing an acacia tree near Mount Kenya, Laikipia, Kenya

1116-52779 - Reticulated giraffe (Giraffa reticulata) crossing an earth dam on the horizon against a blue sky, Laikipia, Kenya

860-291412 - Research assistant working on nitrogen exchanges between bacteria and the roots of legumes in the tropical forest of the "La Selva" research station in Puerto Viejo de Sarapiqui, Costa Rica

489-1807 - Muriwai Gannet Colony, Auckland, North Island, New Zealand, Pacific

1116-51901 - Three African bush elephants (Loxodonta africana) cross rocky pan, Etosha National Park; Otavi, Oshikoto, Namibia

1116-51898 - Two southern giraffes (Giraffa camelopardalis angolensis) stand on dirt track; Otavi, Otjozondjupa, Namibia

1116-51893 - Close-up of head of Southern giraffe (Giraffa giraffa) against a blue sky, Gabus Game Ranch; Otavi, Otjozondjupa, Namibia

1116-51929 - Two women riding horses (Equus ferus caballus) traveling past acacia tree through the bush at the Gabus Game Ranch with mountains in the background at sunset; Otavi, Otjozondjupa, Namibia

1116-51926 - View taken from behind of two women riding horses (Equus ferus caballus) traveling past acacia tree through the bush towards the hills at the Gabus Game Ranch at sunset; Otavi, Otjozondjupa, Namibia

1116-51931 - Two women travelers sitting at a table in the bar talking and drinking at the Gabus Game Ranch; Otavi, Otjozondjupa, Namibia

1116-51905 - Male zebra finch (Taeniopygia guttata) on branch in profile; Otavi, Otjozondjupa, Namibia

1116-51077 - A male cheetah (Acinonyx jubatus) jumps down from the diagonal trunk of a tree. He has brown fur covered with black spots, and in the background can be seen a line of trees, Serengeti National Park; Mara Region, Tanzania

1116-51912 - Female greater kudu (Tragelaphus strepsiceros) stands drinking from waterhole; Otavi, Otjozondjupa, Namibia

1116-51270 - A leopard (Panthera pardus) sits in the forked trunk of a tree. It has a brown, spotted coat and is looking straight at the camera. Shot with a Nikon D850 in Serengeti National Park; Tanzania

1116-51923 - Backlit common warthog (Phacochoerus africanus) walks across sunlit scrub; Otavi, Otjozondjupa, Namibia

1116-51268 - A leopard (Panthera pardus) lies on the branch of a tree with its head up. It has black spots on its brown fur coat and is looking for prey. Shot with a Nikon D850 in the Masai Mara; Kenya

1116-51916 - Common warthog (Phacochoerus africanus) crosses rocky ground in sunshine; Otavi, Otjozondjupa, Namibia

1116-51957 - Woman riding horse (Equus ferus caballus) on a dirt road through the bush on the savanna looking at the camera and smiling at the Gabus Game Ranch; Otavi, Otjozondjupa, Namibia

1116-51949 - Close-up portrait of a plains zebra (Equus quagga - formerly Equus burchellii) eyeing the camera through the bushes in the Etosha National Park; Otavi, Oshikoto, Namibia

1116-51943 - Portrait of a common warthog (Phacochoerus africanus) standing in the bush among rocks and looking at the camera at the Gabus Game Ranch; Otavi, Otjozondjupa, Namibia

1116-51967 - Black rhinoceros and calf (Diceros bicornis) standing in a field of golden long grass on the savanna looking at the camera in Etosh National Park; Otavi, Oshikoto, Namibia

1116-51963 - View taken from behind of two women riding horses following a Southern giraffe (Giraffa camelopardalis angolensis) through the bush at the Gabus Game Ranch; Otavi, Otjozondjupa, Namibia

1116-51266 - A male lion (Panthera leo) shows the Flehmen response by opening his mouth wide as if he were roaring. He has a golden coat that glows in the warm early morning light. Shot with a Nikon D850 in Serengeti National Park; Tanzania

1116-51937 - Overexposed blue wildebeest (Connochaetes taurinus) lying on the Etosha Salt Pan on a sunny day in Etosha National Park; Otavi, Oshikoto, Namibia

1116-51732 - A male leopard (Panthera pardus) stares at the camera. It has a brown, spotted coat, whiskers and a green eye, Masai Mara National Reserve; Kenya

1116-51925 - White Rhinoceros (Ceratotherium simum) walks through the long grass across the savanna in profile in the Etosha National Park at sunset; Otavi, Oshikoto, Namibia

1116-51946 - Close-up of a southern giraffe (Giraffa camelopardalis angolensis) browsing leafy thornbush on the savanna against a blue sky and eyeing the camera at the Gabus Game Ranch; Otavi, Otjozondjupa, Namibia

1116-51952 - Adult Hartmann's mountain zebra (Equus zebra hartmannae) standing in the shade looking forward while foal is looking at the camera at the Gabus Game Ranch at sunrise; Otavi, Otjozondjupa, Namibia

1116-51277 - A brown bear (Ursus arctos) about to catch a salmon in its mouth at the top of Brooks Falls, Alaska. The fish is only a few inches away from its gaping jaws. Shot with a Nikon D800 in Alaska; Kodiak, Alaska, United States of America

1116-51895 - Close-up of head of Southern giraffe (Giraffa giraffa) against a blue sky, Gabus Game Ranch; Otavi, Otjozondjupa, Namibia

1116-51909 - Kirk dik-dik (Madoqua kirkii) walks over sunlit rocky ground; Otavi, Otjozondjupa, Namibia

1116-51918 - Close-up of southern giraffe (Giraffa camelopardalis angolensis) against blurred background; Otavi, Otjozondjupa, Namibia

1116-51941 - Portrait of a gemsbok (Oryx gazella) standing on a grassy plain on the savanna and looking at the camera at the Etosha National Park; Otavi, Oshikoto, Namibia

1116-51939 - Portrait of a sunlit, male common impala (Aepyceros melampus) standing on the the plain casting a long shadow at the Gabus Game Ranch and looking at camera; Otavi, Otjozondjupa, Namibia

1116-51972 - African bush elephant (Loxodonta africana) wading through the mud and drinking from grassy waterhole on the savanna in Etosha National Park looking at camera; Otavi, Oshikoto, Namibia

1116-51271 - An immature black-winged stilt (Himantopus himantopus) with its beak in the water walks through the shallows of a perfectly calm lake, matched by its own reflection. Shot with a Nikon D800 at Ranthambore National Park in India; Rajasthan, India

1116-51942 - Portrait of a gemsbok (Oryx gazella) standing on a grassy plain on the savanna and looking at the camera at the Etosha National Park; Otavi, Oshikoto, Namibia

1116-51962 - Two women riding horses (Equus ferus caballus) on a dirt road traveling through the bush at the Gabus Game Ranch with a southern giraffe (Giraffa camelopardalis angolensis) in the background; Otavi, Otjozondjupa, Namibia

1116-51921 - Black-backed jackal (Canis mesomelas) walks across gravel licking lips, Etosha National Park; Otavi, Oshikoto, Namibia

1116-51267 - A leopard (Panthera pardus) walks towards the camera on a sandy track. It has black spots on its brown fur coat and is raising its paw to take the next step. Shot with a Nikon D850 in the Masai Mara; Kenya

1116-51913 - Female greater kudu (Tragelaphus strepsiceros) standing in tall grass; Otavi, Otjozondjupa, Namibia

1116-51733 - Galapagos giant tortoise (Chelonoidis nigra) lumbers slowly across a long, straight dirt road that stretches off to the horizon. Beyond the grass verge, there is dense forest on either side; Galapagos Islands, Ecuador

1116-51935 - Plains zebra (Equus quagga formerly Equus burchellii) standing in the middle of the long grass on the savanna looking at the camera at the Etosha National Park; Otavi, Oshikoto, Namibia

1116-51907 - Lion (Panthera leo) stands looking ahead on stepping stones, Etosha National Park; Otavi, Oshikoto, Namibia

1116-51953 - Close-up of black rhinoceros (Diceros bicornis) standing beside a rocky waterhole eyeing the camera in the Etosh National Park; Otavi, Oshikoto, Namibia

1116-51965 - Black wildebeest (Connochaetes gnou) standing in the golden long grass of the savanna looking back over shoulder at the camera at the Gabus Game Ranch; Otavi, Otjozondjupa, Namibia

1116-51933 - Portrait of a southern yellow-billed hornbill (Tockus leucomelas) perching in profile on a bush. It has mottled black and brown feathers, a white head and a yellow beak, taken at the Gabus Game Ranch; Otavi, Otjozondjupa, Namibia

1116-51279 - Three Adelie penguins (Pygoscelis adeliae) watch another jumping between two ice floes. They have black heads and backs with white bellies. Shot with a Nikon D810 on Brown Bluff; Antarctica

1116-51924 - Rear view of an African bush elephant (Loxodonta africana) walking away across salt pan in Etosha National Park; Otavi, Oshikoto, Namibia

1116-51964 - Black wildebeest (Connochaetes gnou) standing in the golden long grass of the savanna munching and looking at the camera at the Gabus Game Ranch; Otavi, Otjozondjupa, Namibia

1116-51954 - Close-up of black rhinoceros (Diceros bicornis) standing beside a rocky waterhole eyeing the camera in the Etosh National Park; Otavi, Oshikoto, Namibia

1116-51276 - A Bengal tigress (Panthera tigris tigris) with a catchlight in her eye lies up to her neck in the dark shadows of a water hole. Her name is Maya 'The Enchantress', and she has orange and black stripes with white patches on her head. Shot with a Nikon D810 in Tadoba Andhari Tiger Reserve in India; Chandrapur, Maharashtra, India

1116-51947 - Portrait of male sable antelope (Hippotragus niger) standing in profile at the Gabus Game Ranch at sunset; Otavi, Otjozondjupa, Namibia

1116-51269 - A leopard (Panthera pardus) stands in a tree that is covered in lichen. It has black spots on its brown fur coat and is turning its head to look up. Shot with a Nikon D850 in the Masai Mara; Kenya