832-401370 - Black rhinoceros (Diceros bicornis), Etosha, Namibia, black rhinoceros, Africa

832-401368 - Black rhinoceros (Diceros bicornis), Etosha, Namibia, black rhinoceros, Africa

832-397200 - African elephant (Loxodonta africana) half-headed elephant threat, mammal, close-up in Tsavo East National Park, Kenya, East Africa, Africa

832-397182 - African elephant (Loxodonta africana) a bull threatens, mammal, close-up in Tsavo East National Park, Kenya, East Africa, Africa

832-397183 - African elephant (Loxodonta africana) half-headed elephant threat, mammal, close-up in Tsavo East National Park, Kenya, East Africa, Africa

832-396110 - Bengal tiger (Panthera tigris tigris) Snarling in the water, Tiger Canyon Farm, Philippolis, South Africa, Africa

746-91101 - Southern elephant seal (Mirounga leonina), bull on beach showing threat behaviour. Antarctica, Subantarctica, South Georgia, October

746-91102 - Southern elephant seal (Mirounga leonina), bull on beach showing threat behaviour. Antarctica, Subantarctica, South Georgia, October

1350-2344 - French angelfish, Pomacanthus paru, Ilha rata, Fernando de Noronha national marine sanctuary, Pernambuco, Brazil (S. Atlantic)

1116-46797 - Grey Wolves (Canis Lupus) Demonstrating Hierarchy, Montebello, Quebec, Canada

832-379321 - Two young eagles (Haliaeetus albicilla), fighting on the ground, with dead deer, Masuria, Poland, Europe

869-3602 - gelada baboon portrait male baboon displaying its teeth and gums with lip flipped back threat display Simien Mountains National Park Ethiopia Africa Animals

869-895 - Alaskan brown bear animals close-up of bear showing teeth threat display behaviour danger

999-3 - Mel, the Killer Whale or Orca (Orcinus orca) narrowly misses out on a South American Sea Lion (Otaria flavescens) that it was hunting, in Patagonia. Of the 18 only 7 have mastered the stranding behaviour whereby the Orca enters the shallow surf to feed on Sea Lion pups. Distinctive by his 2 metre dorsal fin, Mel is an expert hunter who feeds on Sea Lion pups before taking them back to his pod. This lucky Sea Lion however managed to escape the hunter's attentions.

999-4 - Mel, the Killer Whale or Orca (Orcinus orca) narrowly misses out on a South American Sea Lion (Otaria flavescens) that it was hunting, in Patagonia. Of the 18 only 7 have mastered the stranding behaviour whereby the Orca enters the shallow surf to feed on Sea Lion pups. Distinctive by his 2 metre dorsal fin, Mel is an expert hunter who feeds on Sea Lion pups before taking them back to his pod. This lucky Sea Lion however managed to escape the hunter's attentions.

999-1 - Mel, the Killer Whale or Orca (Orcinus orca) narrowly misses out on a South American Sea Lion (Otaria flavescens) that it was hunting, in Patagonia. Of the 18 only 7 have mastered the stranding behaviour whereby the Orca enters the shallow surf to feed on Sea Lion pups. Distinctive by his 2 metre dorsal fin, Mel is an expert hunter who feeds on Sea Lion pups before taking them back to his pod. This lucky Sea Lion however managed to escape the hunter's attentions.

999-2 - Mel, the Killer Whale or Orca (Orcinus orca) narrowly misses out on a South American Sea Lion (Otaria flavescens) that it was hunting, in Patagonia. Of the 18 only 7 have mastered the stranding behaviour whereby the Orca enters the shallow surf to feed on Sea Lion pups. Distinctive by his 2 metre dorsal fin, Mel is an expert hunter who feeds on Sea Lion pups before taking them back to his pod. This lucky Sea Lion however managed to escape the hunter's attentions.

817-217622 - Southern Elephant Seal bull mouth wide open in threat or fear display on beach during moulting season, Falkland Islands, January 2003

817-217618 - Southern Elephant Seal bull mouth wide open in threat or fear display on beach during moulting season, Falkland Islands, January 2003

817-217619 - Southern Elephant Seal bull mouth wide open in threat or fear display on beach during moulting season, Falkland Islands, January 2003

817-217646 - Southern Elephant Seal mirounga leonina bull, portrait full face with threat or challange display securing harem during mating season, Island of South Georgia, November 2003